PDFs of publications are provided with the understanding that only one copy will be downloaded per person, and only for personal, academic non-for profit pursuit.
Reports and other publications
Articles and book chapters
130. Wernberg T, Thomsen MS, Baum J, Bishop J, Bruno JF, Coleman MA, Filbee-Dexter K, Gagnon K, He Q, Murdiyarso D, Rogers K, Silliman BR, Smale DA, Starko S, Vanderklift MA. 2024. Impacts of climate change on marine foundation species. Annual Review of Marine Sciences 16: 247–82
129. Hobday A, Burrows M, Filbee-Dexter K, Holbrook N, Sen Gupta A, Smale DA, Smith K, Thomsen MS, Wernberg T. 2023. With the arrival of El Niño, prepare for stronger marine heatwaves. Nature 621:38-41
128. Xu C, Silliman B, Chen J, Li X, Thomsen MS, Zhang Q, Lee J, Lefcheck J, Daleo P, Hughes B, Jones H, Wang R, Wang S, Smith C, Xi X, Altieri A, van de Koppel J, Palmer T, Liu L, Wu J, Li B, He G. 2023. Herbivory limits success of vegetation restoration globally. Science 382: 589-594
127. Cornwall C, Nelson W, Aguirre J, Blain C, Coyle L, D’Archino R, Desmond M, Hepburn C, Liggins L, Shears N, Thomsen MS. 2023. Predicting the impacts of climate change on New Zealand’s seaweed-based ecosystems. New Zealand Journal of Botany: DOI: 10.1080/0028825X.2023.2245786
126. Clemente KJE, Thomsen MS, Zimmerman RC. 2023. The vulnerability and resilience of seagrass ecosystems to marine heatwaves in New Zealand: a remote sensing analysis of seascape metrics using PlanetScope imagery. Remote Sensing in Ecology and Conservation 9: 803–819
125. Montie S, Thomsen MS. 2023. Spatiotemporal stressors, not secondary structures or small temperature increases, control rapid facilitation of intertidal epifauna. Marine Environmental Research 187:105969
124. Montie S, Thoral F, Smith R, Cook F, Tait L, Pinkerton M, Schiel DR, Thomsen MS. 2023. Seasonal trends in marine heatwaves highlight vulnerable coastal ecoregions and historic change points in New Zealand. New Zealand Journal of Marine & Freshwater Research DOI: doi.org/10.1080/00288330.2023.2218102
123. Smith KE, Burrows M, Hobday A, King N, Moore P, Sen Gupta A, Thomsen MS, Wernberg T, Smale D. 2023. Biological Impacts of Marine Heatwaves. Annual Review of Marine Science 15: 119-145
122. Thomsen MS. 2022. Indirect facilitation can unify common interaction chains in Ecology, Zenodo: 1-27. DOI: doi.org/10.5281/zenodo.7455880 PDF
121. Thomsen MS, Altieri A, Angelini C, Bishop MJ, Bulleri F, Farhan R, Frühling V, Gribben P, Harrison S, He Q, Klinghardt M, Langeneck J, Lanham B, Mondardini L, Mulders Y, Oleksyn S, Ramus A, Schiel D, Schneider T, Siciliano A, Silliman B, Smale D, South P, Wernberg T, Zhang S, Zotz G. 2022. Heterogeneity within and among co-occurring foundation species increases biodiversity. Nature communications 13: 581 PDF
120. Alestra T, Bell J, Lewis L, Parsons S, Girvan R, Martin-Paul M, Baggaley F, Blackely T, Moore T, Todd D, Thompson K, Thomsen MS, Hart DE. 2022. Nature based solutions for coastal hazards in Aotearoa New Zealand: results of a nationwide expert survey on the current state of uptake, barriers, and opportunities. Coastal News 5: 62-66 PDF
119. Pullen M, Gerber D, Thomsen MS, Flanagan S. 2022. Seasonal dynamics of faunal diversity and population ecology in an estuarine seagrass bed. Estuaries and Coasts 8: 2578-2591 PDF
118. Thoral F, Montie S, Thomsen MS, Tait LW, Pinkerton M, Schiel DR. 2022. Unravelling seasonal trends in coastal marine heatwave metrics across global biogeographical realms. Scientific reports 12: 7740 PDF
117. Schiel DR, Gerrity S, Orchard S, Alestra T, Dunmore R, Falconer T, Thomsen MS, Tait L. 2021. Cataclysmic disturbances to an intertidal ecosystem: loss of ecological infrastructure slows recovery of biogenic habitats and diversity. Frontiers in Ecology and Evolution: 827 PDF
116. Smith KE, Burrows M, Hobday A, Sen Gupta A, Moore P, Thomsen MS, Wernberg T, Smale D. 2021. Socioeconomic impacts of marine heatwaves – global issues and opportunities. Science 374 (6566): eabj3593 PDF
115. Ren J, Chen J, Xu C, Koppel Jvd, Thomsen MS, Qiu S, Cheng F, Song W, Liu Q, Xu C, Bai J, Zhang Y, Cui B, Bertness MD, Silliman BR, Li B, He Q. 2021. An invasive species erodes the performance of coastal wetland protected areas. Science Advances 7: eabi8943 PDF
114. Thomsen MS, Mondardini L, Thoral F, Gerber D, Montie S, South PM, Tait L, Schiel DR. 2021 Cascading impacts of earthquakes and extreme heatwaves have destroyed populations of an iconic marine foundation species. Diversity and Distributions 27: 2369-2383 PDF
113. Bass A, Wernberg T, Thomsen MS, Smale DS. 2021. Another decade of marine climate change experiments: trends, progress and knowledge gaps. Frontiers in Marine Science 8: 1233 PDF
112. Thomsen MS, Montie S, Gerber D, Thoral F, South PM, Tait L, Wernberg T, Smale D, Schiel DR. 2021. Impacts of marine heatwaves: what are they, what have they done and what can we expect in the future? Coastal News 76: 3-6 PDF
111. Tait LW, Thoral F, Pinkerton MH, Thomsen MS, Schiel DR. 2021. Loss of giant kelp, Macrocystis pyrifera, driven by marine heatwaves and exacerbated by poor water clarity in New Zealand. Frontiers in Marine Science 8: 1168 PDF
110. Schiel DR, Alestra A, Gerrity S, Orchard S, Dunmore R, Crossett D, Thomsen MS, Pirker J, Tait L, 2020. Legacies of the Kaikōura earthquake on the coastal marine ecosystem. Coastal News 72: 3-5 PDF
109. Wernberg T. Couraudon-Réale M, Thomsen MS, Tuya F, 2020. Kelp understory communities are modulated by gradients in disturbance intensity, extent and ocean climate. Marine Ecology Progress Series 651: 57-69 PDF
108. Gupta AS, Thomsen MS, Benthuysen JA, Hobday AJ, Oliver E, Alexander, LV, Burrows MT, Donat, MG, Feng M, Holbrook NJ, Perkins-Kirkpatrick S, Moore PJ, Rodrigues RR, Scannell HA, Taschetto AS, Ummenhofer CC, Wernberg T, Smale DA. 2020. Drivers and impacts of the most extreme marine heatwaves events. Scientific reports 10: 1-15 PDF
107. Salinger JM, Diamond H, Behrens E, Fernandez D, Fitzharris B, Herold M. Johnstone P, Kerckhoffs H, Mullan B, Parker A, Renwick J, Schofield C, Siano A, Smith R, South P, Sutton PJ, Teixeira E, Thomsen MS, Trought M. 2020. Unparalleled coupled ocean-atmosphere summer heatwaves in the New Zealand region: drivers, mechanisms and impacts. Climatic Change 162: 485–506 PDF
106. Montie S, Thomsen MS, Rack W, Broady PA. 2020. Extreme summer marine heatwaves increase chlorophyll a in the Southern Ocean. Antarctic Science 32: 508-509 PDF
105. Gribben PE, Poore AGB, Thomsen MS, Quesey P, Weschke E, Wright JT. 2020. Habitat provided by native species facilitates higher abundances of an invader in its introduced compared to native range. Scientific reports 10: 1-9 PDF
104. Thomsen MS, Metcalfe I, Siciliano A, South PM, Gerrity S, Alestra T, Schiel DR. 2020. Earthquake-driven destruction of an intertidal habitat cascade. Aquatic Botany 164, 103217 PDF
103. Thomsen MS. 2020. Indiscriminate data aggregation in ecological meta-analysis underestimates impacts of invasive species. Nature Ecology and Evolution 4: 313-314 PDF
102. Staehr PA, Jakobsen HH, Hansen JLS, Andersen P,Christensen J, Göke C, Thomsen MS. 2020. Trends in records and contribution of non-indigenous species to marine communities in Danish waters. Aquatic Invasions 15: 217-244 PDF
101. Vieira, A, Martin A, Engelen AH, Thomsen MS, Arenas F. 2020. Interactive effects of co-occurring anthropogenic stressors on the seagrass, Zostera noltei. Ecological Indicators 109: 105780 PDF
100. Oliver EC, Burrows MT, Donat MG, Sen Gupta A, Alexander LV, Perkins-Kirkpatrick SE, Benthuysen J, Hobday AJ, Holbrook NJ, Moore PJ, Thomsen MS. 2019. Projected marine heatwaves in the 21st century and the potential for ecological impact. Frontiers in Marine Science 6: 734 PDF
99. Straub S, Wernberg T, Thomsen MS, Moore PJ, Burrows M, Harvey BP. Smale DA. 2019. Resistance to obliteration; responses of seaweeds to marine heatwaves. Frontiers in Marine Science 6: 763 PDF
98. Wernberg T, Coleman MA, Bell S, Babcock RC, Bolton JJ, Connell SD, Hurd CL, Johnson CR, Marzinelli E, Shears NT, Steinberg PD, Thomsen MS, Vanderklift MA, Verges A, Wright JT. 2019. Biology and ecology of the globally significant kelp Ecklonia radiata. Oceanography and Marine Biology: An Annual Review 57: 263-136 PDF
97. Gribben PE, Angelini C, Altieri AH, Bishop MJ, Thomsen MS, Bulleri F. 2019. Facilitation cascades in marine ecosystems: A synthesis and future directions. Oceanography and Marine Biology: An Annual Review 57: 95-136 PDF
96. Siciliano A, Schiel DR, Thomsen MS. 2019. Effects of local anthropogenic stressors on a habitat cascade in an estuarine seagrass system. Marine and Freshwater Research 70: 1129-1142 PDF
95. Schiel DR, Alestra T, Gerrity S, Orchard S, Dunmore R, Pirker J, Lilley SA, Tait L, Thomsen MS. 2019. The Kaikōura earthquake in southern New Zealand: loss of connectivity of marine communities and the necessity of a cross-ecosystems perspective. Aquatic Conservation: Marine and Freshwater Ecosystems 29: 1520-1534 PDF
94. Thomsen MS, South PM. 2019. Communities and attachment networks associated with primary, secondary and alternative foundation species; a case of stressed and disturbed stands of Southern Bull Kelp. 2019. Diversity 11(4): 56 PDF
93. Thomsen MS, Gerrity S, Alestra T, South PM, Lilley S, Monstrani L, Schiel DR. 2019. Local extinction of bull kelp (Durvillaea spp.) due to a marine heatwave. 2019. Frontiers Marine Science 6: 84 PDF
92. Smale D, Wernberg T, Oliver E, Thomsen MS, Harvey B, Straub S, Burrows M, Alexander L, Benthuysen J, Donat M, Feng M, Hobday A, Holbrook N, Perkins-Kirkpatrick S, Scannell H, Sen Gupta A, Payne B, Moore P. 2019. Marine heatwaves threaten global biodiversity and the provision of ecosystem services. Nature Climate Changes 9: 306-312 PDF
91. Thomsen MS, Ramus AP, Long ZT, Silliman BR. 2019. A seaweed increases ecosystem multifunctionality when invading bare mudflats. Biological Invasions 21: 27-36 PDF
90. Thomsen MS, Alestra T, Brockerhoff D, Lilley SA, South PM, Schiel DR. 2018. Modifications of kelp seasonality and invertebrate diversity where an invasive kelp co-occurs with native mussels. Marine Biology 165: 173 PDF
89. Martínez B, Wernberg T, Radford B, Thomsen MS, Russell BD, Connell SC, Bradshaw CJA, Carreño F, Fordham DA, Gurgel CFD. 2018. Distribution models predict large contractions in habitat-forming seaweeds in response to ocean warming. Diversity and Distributions 24: 1350-1366 PDF
88. Hobday AJ, Oliver ECJ, Gupta AS, Benthuysen JA, Burrows MT, Donat MG, Holbrook NJ, Moore PJ, Thomsen MS, Wernberg T, Smale DA. 2018. Categorizing and Naming Marine Heatwaves. Oceanography. 31: 63-73 PDF
87. Thomsen MS, Altieri AH, Angelini C, Bishop MJ, Gribben PE, Lear G, He Q, Schiel DR, Silliman BR, South PM, Watson DM, Wernberg T, Zotz G. 2018. Secondary Foundation Species Enhance Biodiversity. Nature Ecology and Evolution 2: 634-639 PDF
86. Schiel DR, Gerrity S, Alestra T, Pirker J, Marsden I, Dunmore R, Tait L, South PM, Taylor D, Thomsen MS. 2018. Kaikōura earthquake: summary of impacts and changes in nearshore marine communities. In: Shaky Shores – Coastal impacts & responses to the 2016 Kaikōura earthquakes”. NZCS Special Publications series v. 3 p 25-28 PDF
85. Wernberg T, Coleman MA, Bennett S, Thomsen MS, Tuya F, Kelaher BP. 2018. Genetic diversity and kelp forest vulnerability to climatic stress. Scientific Reports 8:1851 PDF
84. Ramus AP. Silliman BR, Thomsen MS, Long ZT. 2017. An invasive foundation species enhances multifunctionality in a coastal ecosystem. Proceedings of the National Academy of Sciences 114: 8580-8585 PDF
83. South PM, Floerl O, Forrest BM, Thomsen MS. 2017. A review of three decades of research on the invasive kelp Undaria pinnatifida in Australasia: An assessment of its success, impacts and status as one of the world’s worst invaders. Marine environmental research 131:243-257 PDF
82. Thomsen MS, Hildebrand T, South PM, Foster T, Siciliano A, Oldach E, Schiel DR. 2016. A sixth-level habitat cascade increases biodiversity in an intertidal estuary. Ecology and Evolution 6: 8291–8303 PDF
81. Wernberg T. Bennett S., Babcock RC, Bettignies Td, Cure K, Depczynski M, Dufois F, Fromont J, Fulton JC, Hovey RK, Harvey ES, Holmes TH, Kendrick GA, Radford B, Santana-Garcon J, Saunders BJ, Smale DA, Thomsen MS, Tuckett CA, Tuya F, Vanderklift MA, Wilson S. 2016. Climate driven regime shift of a temperate marine ecosystem. Science 353: 169-172 PDF
80. South PM, Thomsen MS. 2016. The ecological role of invading Undaria pinnatifida: an experimental test of the driver-passenger models. Marine Biology 163: 1-15 PDF
79. Thomsen MS, Wernberg T, South PM, Schiel DR. 2016. To include or not to include (the invader in community analyses)? That is the question. Biological Invasions 18: 1515-152 PDF.
78. Thomsen MS, Wernberg T, Stæhr T, Schiel D. 2016. Ecological Interactions between Marine Plants and Alien Species. In: Marine Macrophytes as Foundation Species (Edi. Ólafsson E), Science Publisher/CRC Press. 226-258. ISBN 9781498723244 PDF
77. Thornber CS, Jones E, Thomsen MS. Epibiont-Marine Macrophyte Assemblages. In: Marine Macrophytes as Foundation Species (Edi. Ólafsson E), Science Publisher/CRC Press, p. 43-75. ISBN 9781498723244 PDF
76. Wernberg T, Arenas F, Olabarria C, Thomsen MS, Mohring MB. 2016. Threats to Ecosystem Engineering Macrophytes: Climate Change. In: Marine Macrophytes as Foundation Species (Edi. Ólafsson E), Science Publisher/CRC Press. p. 201-225 ISBN 9781498723244 PDF
75. Thomsen MS, Wernberg T, South P, Schiel DR. 2016. Non-native seaweeds drive changes in marine coastal communities around the world. In ‘Seaweed Phylogeography‘ (Edi. Ceridwen Fraser and Hu Zi-Min) Springer Press. p. 147-185. ISBN 978-94-017-7534-2 PDF
74. Straub, S., Wernberg, T. Thomsen MS. 2016. Seaweed range expansions. Book chapter to ‘Seaweed Phylogeography’ In ‘Seaweed Phylogeography‘ (Edi. Ceridwen Fraser and Hu Zi-Min) Springer Press. p. 63-93. ISBN 978-94-017-7534-2 PDF
73. South P, Lilley S, Tait L, Alestra T, Hickford M, Thomsen MS, Schiel D. 2016. Transient effects of an invasive kelp on the community structure and primary productivity of an intertidal assemblage. Marine and Freshwater Research 67: 103-112. DOI: http://dx.doi.org/10.1071/MF14215 PDF
72. Thomsen MS, Metcalfe I, South P, Schiel DR. 2016. A host-specific habitat former controls biodiversity across ecological transitions in a rocky intertidal facilitation cascade. Marine and Freshwater Research 67: 144-152. DOI: http://dx.doi.org/10.1071/MF14152 PDF
71. Tait LW, South P, Lilley SA, Thomsen MS, Schiel DR. 2015. Assemblage and understory carbon production of native and invasive canopy forming macroalgae. Journal of Experimental Marine Biology and Ecology 469: 10–17. DOI: http://dx.doi.org/10.1016/j.jembe.2015.04.007 PDF
70. Thyrring J, Thomsen MS, Brunbjerg AK, Wernberg T. 2015. Diversity and abundance of epibiota on invasive and native estuarine gastropods depend on substratum and salinity. Marine and Freshwater Research 66: 1-10. DOI: http://dx.doi.org/10.1071/MF14311. PDF
69. Thomsen MS, Wernberg T. 2015. The devil in the detail: harmful seaweeds are not harmful to everyone. Global Change Biology 21: 1381–1382. DOI:10.3354/meps10566 PDF Online Supplement
68. Thomsen MS, Wernberg T, Schiel D. 2015. Impacts of Marine Invasions on Biodiversity and Ecosystem Functioning. In: ‘Marine ecosystems: human impacts on biodiversity, functioning and services‘ (Edi. Crowe, Frid, & Austen), Cambridge University Press’s Ecology, Biodiversity and Conservation series p. 274-332, ISBN: 9781107037670 PDF
67. Wernberg T, Russell BD, Thomsen MS, Connell SD. 2014. Marine biodiversity and climate change. In: ‘Global Environmental Change, Handbook of Global Environmental Pollution‘ (Edi. Freedman B.) Vol. 1. Springer-Verlag Berlin Heidelberg. p. 181-187. DOI: 10.1007/978-94007-5784-4_80 PDF
66. Thomsen MS, Wernberg T, Olden JD, Byers JE, Bruno JF, Silliman BR, Schiel DR. 2014. Forty years of experiments on invasive species: are biases limiting our understanding of impacts? Neobiota 22: 1-22. DOI: 10.3897/neobiota.22.6224 PDF Online Supplement
65. Thomsen MS, Byers JE, Schiel DR, Bruno JF, Olden JD, Wernberg T, Silliman BR. 2014. Impacts of marine invaders on biodiversity depend on trophic position and functional similarity. Marine Ecology Progress Series 495: 39–47. DOI: 10.3354/meps10566 PDF Online Supplement
64. Thomsen MS, Wernberg T. 2014. On the generality of cascading habitat formation. Proc. Roy. Soc. London Ser B 281(1777):20131994. DOI: 10.1098/rspb.2013.1994. PDF Online Supplement
63. Wernberg T, Russell BD, Bradshaw CJA, Gurgel CFD, Thomsen MS, Poloczanska ES, Connell SD. 2014. Misconceptions about analyses of Australian seaweed collections. Phycologia 53: 215-220. DOI: http://dx.doi.org/10.2216/13-197.1 PDF
62. Gurgel CFD, Wernberg T, Thomsen MS, Russell BD, Adam, P, Waters MJ, Connell SD. 2014 Shared patterns of species turnover between seaweeds and seed plants break down at increasing distances from the sea. Ecology and Evolution 4: 27-34, DOI: 10.1002/ece3.893. PDF
61. Thyrring J, Thomsen MS, Wernberg T. 2013. Large scale facilitation of a sessile community by an invasive habitat-forming snail. Helgolander Marine Research 67: 789-794. DOI: 007/s10152-013-0363-2 PDF Online Supplement
60. Thomsen MS, Staehr PA, Nejrup LB, Schiel DR. 2013. Effects of the invasive macroalga Gracilaria vermiculophylla on two co-occurring foundation species and associated invertebrates. Aquatic Invasions 8: 133-145. DOI: http://dx.doi.org/10.3391/ai.2013.8.2.02 PDF
59. Wernberg T, Thomsen MS, Connell SD, Russell BD, Waters JM, Zuccarello GC, Kraft GT, Sanderson C, West JA, Gurgel CFD. 2013. The Footprint of Continental-Scale Ocean Currents on the Biogeography of Seaweeds. PLOS one 8:e80168. DOI: 10.1371/journal.pone.0080168 PDF
58. Wernberg T, Thomsen MS, Kotta J. 2013. Complex plant–herbivore–predator interactions in a brackish water seaweed habitat. Journal of Experimental Marine Biology and Ecology 449: 51-56. DOI: http://dx.doi.org/10.1016/j.jembe.2013.08.014 PDF
57. Nejrup LB, Staehr PA, Thomsen MS. 2013. Temperature and light dependent growth and metabolism of the invasive red algae Gracilaria vermiculophylla (Ohmi) Pappenfuss – a comparative study with two native macroalgae. European Journal of Phycology 48: 295–308. DOI: http://dx.doi.org/10.1080/09670262.2013.830778 PDF
56. Wernberg T, Smale DA, Tuya F, Thomsen MS, Langlois TJ, De Bettignies T, Bennett S, Rousseaux CS. 2013. An extreme climatic event alters marine ecosystem structure in a global biodiversity hotspot. Nature Climate Change 3: 78-82. DOI: 10.1038/nclimate1627 PDF Online Supplement
55. De Bettignies T, Thomsen MS, Wernberg T. 2012. Wounded kelps: patterns and susceptibility to breakage. Aquatic Biology 17: 223-233. DOI 10.3354/ab00471 PDF
54. Thomsen MS, de Bettignies T, Wernberg T, Holmer M, Debeuf B. 2012. Harmful algae are not harmful to everyone. Harmful Alga 16: 74-80. DOI: 10.1016/j.hal.2012.01.005 PDF
53. Wernberg T, Smale DA, Thomsen MS. 2012. A decade of climate change experiments on marine organisms: procedures, patterns and problems. Global Ecology Biology 18: 1491-1498. DOI: 10.1111/j.1365-2486.2012.02656.x PDF Online Supplement
52. Hoeffle H, Wernberg T, Thomsen MS, Holmer M. 2012. Drift algae, an invasive snail and elevated temperature reduces the ecological performance of a warm-temperate seagrass, through additive effects. Marine Ecology Progress Series 450: 67-80. DOI: 10.3354/meps09552 PDF
51. McCoy MW, Altieri AH, Holdredge C, Thomsen MS, Silliman BR. 2012. Facilitation. In: ‘Encyclopedia of theoretical ecology‘ (Edi. Hastings A, Gross LJ), University of California Press. p. 276-280. ISBN: 9780520269651 PDF
50. Thomsen MS, Wernberg T, Engelen AH, Tuya F, Vanderklift MA, Holmer M, McGlathery KJ, Arenas F, Kotta J, Silliman BR. 2012. A meta-analysis of seaweeds impact on seagrasses: generalities and knowledge gaps. PLoS One 7: e28595. DOI: 10.1371/journal.pone.0028595 PDF Online Supplement
49. Ramsey E III, Rangoonwala A, Thomsen MS, Schwarzschild A. 2012. Flat-plate techniques for measuring reflectance of macro-algae (Ulva curvata). International Journal of Remote Sensing 33: 3147-3155. DOI: 10.1080/01431161.2011.633124 PDF
48. Ramsey E III, Rangoonwala A, Thomsen MS, Schwarzschild A. 2012. Spectral definition of the macro-algae Ulva curvata in the back-barrier bays of the Eastern Shore of Virginia, USA. International Journal of Remote Sensing 33: 586-603. DOI: 10.1080/01431161.2010.543436 PDF
47. Wernberg T, Russell B, Thomsen MS, Gurgel CFD, Bradshaw CJA, Poloczanska ES, Connell SD. 2011. Seaweed communities in retreat from ocean warming. Current Biology 21: 1828-1832. DOI: 10.1016/j.cub.2011.09.028 PDF Online Supplement
46. Thomsen MS, Ferdie M, McGlathery KJ. 2011. Tube-forming polychaetes enhance invertebrate diversity and abundance in sandy sediments of Mozambique, Africa. African Journal of Marine Science 33: 327–332. DOI: 10.2989/1814232X.2011.600433 PDF
45. Thomsen MS, Olden J, Wernberg T, Griffin J, Silliman BR. 2011. A broad framework to organize and compare invasion impacts. Environmental Research 111: 899 –908. DOI: 10.1016/j.envres.2011.05.024 PDF Online Supplement
44. Tuya F, Wernberg T, Thomsen MS. 2011. The relative influence of local to regional drivers of variation in reef fishes. Journal of Fish Biology 79: 217-234. DOI: 10.1111/j.1095-8649.2011.03015.x PDF
43. Tuya F, Vanderklift MA, Wernberg T, Thomsen MS. 2011. Gradients in the number of species at reef-aeagrass ecotones explained by gradients in abundance. PLoS One 6: e20190. DOI: 10.1371/journal.pone.0020190 PDF
42. Thomsen MS, Wernberg T, Olden J, Griffin J, Silliman BR. 2011. A general framework to address context-dependent invasion impact. Journal of Experimental Marine Biology and Ecology 400: 322-327. DOI: 10.1016/j.jembe.2011.02.033 PDF
41. Wernberg T, Thomsen MS, Tuya F, Kendrick G. 2011. Biogenic habitat structure on reefs change along a latitudinal gradient in temperate ocean climate. Journal of Experimental Marine Biology and Ecology 400: 264-271. DOI: 10.1016/j.jembe.2011.02.017 PDF
40. Hoeffle H, Thomsen MS, Holmer M. 2011. High mortality of Zostera marina under high temperature regimes but minor effects of the invasive macroalgae Gracilaria vermiculophylla. Estuarine Coastal and Shelf Science 92: 35-46. DOI: 10.1016/j.ecss.2010.12.017 PDF
39. Holmer M, Wirachwong P, Thomsen MS. 2011. Negative effects of stress-resistant drift algae and high temperature on a small ephemeral seagrass species. Marine Biology 158: 297-309. DOI: 10.1007/s00227-010-1559-5 PDF
38. Waters JM, Wernberg T, Connell SD, Thomsen MS, Zuccarello J, Kraft GT, Sanderson C, West JA, Gurgel CFD. 2010. Australia’s marine biogeography revisited: back to the future? Austral Ecology 35: 988–992. DOI: 10.1111/j.1442-9993.2010.02114.x PDF
37. Wernberg T, Tuya F, Thomsen MS, Kendrick GA. 2010. Turban snails as habitat for foliose algae: contrasting geographical patterns in species richness. Marine & Freshwater Research 61: 1-6. DOI: 10.1071/MF09184 PDF
36. Thomsen MS. 2010. Experimental evidence for positive effects of invasive seaweed on native invertebrates via habitat-formation in a seagrass bed. Aquatic Invasions 5: 1-6. DOI: 10.3391/ai.2010.5.4.02 PDF
35. Thomsen MS, Altieri A., Wernberg T, Tuya F, Gulbransen D, McGlathery K, Holmer M, Silliman BR. 2010. Habitat cascades: the conceptual context and global relevance of facilitation cascades via habitat formation and modification. Integrative and Comparative Biology 50: 158-175. DOI:10.1093/icb/icq042 PDF Online Supplement
34. Tuya F, Vanderklift M, Wernberg T, Thomsen MS, Hynes G, Hanson C. 2010. Proximity to edges impacts cross-habitat flows: implications for abundance patterns and species interactions. Marine Ecology Progress Series 405: 175–186. DOI: 10.3354/meps08516 PDF
33. Wernberg T, Thomsen MS, Tuya F, Kendrick G, Staerh P. 2010. The resilience of Australasian kelp beds decrease along a latitudinal gradient in ocean temperature. Ecology Letters 13: 685–694. DOI: 10.1111/j.1461-0248.2010.01466.x PDF
32. Thomsen MS, Wernberg T, Tuya F, Silliman BR. 2010. Ecological performance and possible origin of a ubiquitous but under-studied gastropod. Estuaries, Coastal and Shelf Sciences 87: 501-650. DOI: 10.1016/j.ecss.2010.02.014 PDF Online Supplement
31. Thomsen MS, Adam P, Silliman BR. 2009. Anthropogenic threats to Australasian coastal salt marshes. In: ‘Human Impacts on Salt Marshes: A Global Perspective‘ (Edi. Silliman BR, Bertness MD, Strong D), University of California Press. p. 361-390. ISBN: 9780520258921 PDF
30. Silliman BR, Bertness MD, Thomsen MS. 2009. Trophic cascades and climate change in southern US marshes: Will the over-harvesting of blue crabs and drought stress trigger massive die-off of southern salt marshes? In: ‘Human Impacts on Salt Marshes: A Global Perspective‘ (Edi. Silliman BR, Bertness MD, Strong D), University of California Press. p. 103-114. ISBN: 9780520258921 PDF
29. Thomsen MS, McGlathery KJ, Schartschild A, Silliman BR. 2009. Distribution and ecological role of the non-native macroalgae Gracilaria vermiculophylla in Virginia salt marshes. Biological Invasions 11: 2303-2316. DOI: 10.1007/s10530-008-9417-9 PDF
28. Tuya F, Wernberg T, Thomsen MS. 2009. Habitat structure affect abundances of labrid fishes across temperate reefs in south-western Australia. Environmental Biology of Fishes. 86: 311–319. DOI: 10.1007/s10641-009-9520-5 PDF
27. Thomsen MS, Wernberg B, Silliman BR, Tuya F. 2009. Evidence for impacts of non-indigenous macroalgae: a meta-analysis of experimental field studies. Journal of Phycology 45: 10-15. DOI: 10.1111/j.1529-8817.2009.00709.x PDF Online Supplement
26. Nyberg C, Thomsen MS, Wallentinus I. 2009. Flora and fauna associated with the introduced red alga Gracilaria vermiculophylla. European Journal of Phycology. 44: 395-403. DOI: 10.1080/09670260802592808 PDF
25. Thomsen MS, Wernberg T, Silliman BR, Josefson A. 2009. Broad-scale patterns of abundance of non-indigenous soft-bottom invertebrates in Denmark. Helgoland Marine Research 63: 159-167. DOI: 10.1007/s10152-008-0142-7 PDF
24. Tuya F, Wernberg T, Thomsen MS. 2009. Colonization of gastropods on subtidal reefs depends on density in adjacent habitats, not on disturbance regime. Journal of Molluscan Studies 75: 27–33. DOI: 10.1093/mollus/eyn032 PDF
23. Tuya F, Wernberg T, Thomsen MS. 2008. Testing the ‘abundant centre’ hypothesis on endemic reef fishes in south-western Australia. Marine Ecology Progress Series 372: 225–230. DOI: 10.3354/meps07718 PDF
22. Thomsen MS, Wernberg T, Stæhr PA, Silliman BR, Josefson AB, Krause-Jensen D, Risgaard-Petersen N. 2008. Annual changes in abundance of non-indigenous marine benthos on a very large spatial scale. Aquatic Invasions 3: 133-139. DOI: 10.3391/ai.2008.3.2.3 PDF
21. Tuya F, Wernberg T, Thomsen MS. 2008. The spatial arrangement of reefs alters the ecological patterns of fauna between interspersed algal habitats. Estuarine, Coastal and Shelf Science 78: 774–782. DOI: 10.1016/j.ecss.20 08.02.017 PDF
20. Thomsen MS, McGlathery KJ. 2007. Stress tolerance of the invasive macroalgae Codium fragile and Gracilaria vermiculophylla in a soft-bottom turbid lagoon. Biological Invasions 9: 1-12. DOI: 10.1007/s10530-006-9043-3 PDF
19. Thomsen MS, Stæhr P, Nyberg C, Krause-Jensen D, Schwærter S, Silliman BR. 2007. Gracilaria vermiculophylla in northern Europe, with focus on Denmark, and what to expect in the future. Aquatic Invasions 2: 83-94. DOI: 10.3391/ai.2007.2.2.1 PDF
18. Thomsen MS, Wernberg T, Stæhr PA, Krause-Jensen D, Risgaard-Petersen N, Silliman BR. 2007. Alien macroalgae in Denmark – a broad-scale national perspective. Marine Biology Research 3: 61 – 72. DOI: 10.1080/17451000701213413 PDF
17. Thomsen MS, Silliman BR, McGlathery KJ. 2007. Spatial variation in recruitment of native and invasive sessile species onto oyster reefs in a temperate soft-bottom lagoon. Estuarine, Coastal & Shelf Science 72: 89-101. DOI: 10.1016/j.ecss.2006.10.004 PDF
16. Thomsen MS, McGlathery KJ, Tyler AC. 2006. Macroalgal distribution pattern in a shallow, soft-bottom lagoon, with emphasis on the nonnative Gracilaria vermiculophylla and Codium fragile. Estuaries and Coasts 29: 465-473. DOI: 10.1007/BF02784994 PDF
15. Thomsen MS, Wernberg T, Stæhr PA, Pedersen MF. 2006. Spatio-temporal distribution patterns of the invasive macroalga Sargassum muticum within a Danish Sargassum-bed. Helgoland Marine Research 60: 50-58. DOI: 10.1007/s10152-005-0016-1 PDF
14. Thomsen MS, McGlathery KJ. 2006. Effects of accumulations of sediments and drift algae on recruitment of sessile organisms associated with oyster reefs. Journal of Experimental Marine Biology and Ecology 328: 22-34. DOI: 10.1016/j.jembe.2005.06.016 PDF
13. Thomsen MS, Gurgel CFD, Fredericq S, McGlathery KJ. 2006. Gracilaria vermiculophylla (Rhodophyta, Gracilariales) in Hog Island Bay, Virginia: a cryptic alien and invasive macroalgae and taxonomic corrections. Journal of Phycology 42: 139-141. DOI: 10.1111/j.1529-8817.2006.00160.x PDF
12. Thomsen MS, Wernberg T. 2005. Miniview: What affects the forces required to break or dislodge macroalgae? European Journal of Phycology 40: 1-10. DOI: 10.1080/09670260500123591 PDF
11. Thomsen MS, McGlathery K. 2005. Facilitation of macroalgae by the sedimentary tube forming polychaete Diopatra cuprea. Estuarine, Coastal and Shelf Science 62: 63-73. DOI: 10.1016/j.ecss.2004.08.007 PDF
10. Thomsen MS, Krause-Jensen D, Wernberg T, Stæhr PA, Risgaard NP. 2005. Alien seaweeds in Denmark: Where? When? How many? (in Danish). Urt 29: 110-115 PDF
9. Wernberg T, Thomsen MS. 2005. The effect of wave exposure on the morphology of Ecklonia radiata in southwestern Australia. Aquatic Botany 83: 61-70. DOI: 10.1016/j.aquabot.2005.05.007 PDF
8. Pedersen MF, Stæhr PA, Wernberg T, Thomsen MS. 2005. Biomass dynamics of exotic Sargassum muticum and native Halidrys siliquosa in Limfjorden, Denmark – implications of species replacements on turnover rates. Aquatic Botany 83: 31-47. DOI: 10.1016/j.aquabot.2005.05.004 PDF
7. Thomsen MS, Wernberg T, Kendrick G. 2004. The effect of thallus size, life stage, aggregation, wave exposure and substrate conditions on the forces required to break or dislodge the small kelp Ecklonia radiata. Botanica Marina 47: 454-460. DOI: 10.1515/BOT.2004.068 PDF
6. Thomsen MS 2004. Species, thallus size and substrate determine macroalgal break forces and break places in a low energy soft bottom lagoon. Aquatic Botany 80: 153-161. DOI: 10.1016/j.aquabot.2004.08.002 PDF
5. Wernberg T, Thomsen MS, Stæhr PA, Pedersen MF. 2004. Epibiota communities of the introduced and indigenous macroalgal relatives Sargassum muticum and Halidrys siliquosa in Limfjorden (Denmark). Helgoland Marine Research 58: 154-161. DOI: 10.1007/s10152-004-0180-8 PDF
4. Wernberg T, Coleman M, Fairhead A, Miller S, Thomsen MS. 2003. Morphology of Ecklonia radiata (C. Ag.) J. Agardh. along its geographic distribution in Southwestern Australia and Australasia. Marine Biology 143: 47-55. DOI: 10.1007/s00227-003-1069-9 PDF
3. Wernberg T, Thomsen MS, Stæhr PA, Pedersen MF. 2001. Comparative phenology of Sargassum muticum and Halidrys siliquosa (Phaeophycae: Fucales) in Limfjorden, Denmark. Botanica Marina 44: 31-49. DOI: 10.1515/BOT.2001.005 PDF
2. Stæhr PA, Pedersen MF, Thomsen MS, Wernberg T, Krause-Jensen D. 2000. The Invasion of Sargassum muticum in Limfjorden (Denmark) and its possible impact on the indigenous macroalgal community. Marine Ecology Progress Series 207: 79-88. DOI: 10.3354/meps207079 PDF
1. Wernberg T, Thomsen MS, Stæhr PA. (1998). The invasion of japweed (Sargassum muticum) in Denmark – status anno 1998 (in Danish). Urt 4: 128-132 PDF
Reports and other publications
14. Smale D, Thomsen MS, Bennett S, Duarte CM, Eger A, Filbee-Dexter K, Flack B, Gundersen H, Hancke K, Hepburn C, Jackson A, King N, Lavoie C, Moore P, Norderhaug K, Schoenrock K, Smit AJ, Vásquez JA, Watanabe A, Wernberg T. 2023. Biodiversity and ecosystem services. United Nations Environment Programme. Into the Blue: Securing a Sustainable Future for Kelp Forests. Nairobi: 45-70.
13. Wernberg T, Buschmann A, Krumhansl KA, Cavanaugh K, Thomsen MS, Smale D. 2023. Ecology of kelp forests and their main threats. United Nations Environment Programme. Into the Blue: Securing a Sustainable Future for Kelp Forests. Nairobi:9-27
12. Sen Gupta A, Benthuysen J, Burrows M, Filbee-Dexter K, Frolicher T, Hobday A, Holbrook N, Moore P, Oliver E, Smale D, Smith K, Thomsen MS, Wernberg T. 2023. Marine heatwaves: definition duel heats up. Nature 617: 465-465
11. Thomsen MS, Moser A, Pullen M, Gerber D, Flanagan S. 2020. Seagrass beds provide habitats for crabs, shrimps and fish in two estuaries on the South Island of New Zealand. Report for Environment Canterbury. Pp. 20 PDF
10. Thomsen MS, Oldach E. 2016. Habitat cascades. Wikipedia https://en.wikipedia.org/wiki/Habitat_cascade
9. Hansen JW, Stæhr PA, Mohn C, Thomsen MS. 2012. Evaluation of ballast water risk assessment. Research note from DCE – Danish Centre for Environment and Energy. Pp. 45 PDF
8. Thomsen MS, Stæhr PA. 2012. Third Symposium on Danish Marine Bioinvasions, Roskilde: PDF Abstract Book. Pp. 14 PDF
7. Stæhr PA, Thomsen MS. 2011. Notatudkast – Opgørelse over rumlig udbredelse, tidslig udvikling og tæthed af introducerede arter i danske farvande (Reviews; Spatio-temporal disribution and densities of introduced marine species in Danish waters). Report for Danish National Environmental Research Academy. Pp. 21 PDF
6. Thomsen MS, Stæhr PA. 2010. Second Symposium on Danish Marine Bioinvasions, Copenhagen: PDF Abstract Book – With Introduction & Summery. Pp. 19 PDF
5. Thomsen MS, Wernberg T. 2009. Drift algae, invasive snails and seagrass health in the Swan River: patterns, impacts and linkages. Final report the Swan River Trust, Report no 2009-2 from the Centre for Marine Ecosystems Research, Edith Cowan University, Perth, Australia. Pp.105 PDF
4. Wernberg T, Thomsen MS, McMahon K. 2008. Determining seagrass depth distribution in the Swan River – a test of the dumpy level method with recommendations for alternatives. Report prepared for the Water Science Branch, Department of Water. Report no 2008-2 from the Centre for Marine Ecosystems Research, Edith Cowan University, Perth, Australia. Pp. 21 PDF
3. Thomsen MS, Wernberg T, Stæhr PA, Krause-Jensen D, Josefson A, Tendal O. 2008. Introduced animals and plants in Denmark – terminology, mechanisms and effects (in Danish). Naturens Verden 6:10-18 PDF
2. Thomsen MS, Stæhr PA, Wernberg T. 2007. First Symposium on Danish Marine Bioinvasions, Copenhagen: PDF Abstract Book – With Introduction & Summery. Pp. 18 PDF
1. Thomsen MS, Josefson A, Wernberg T. 2007. Introduced benthic marine invertebrates: how abundant are they in Denmark? (In Danish) Vand og Jord 2: 44-48 PDF